Retroviral Immunology
Quentin J Sattentau
Professor in Immunology
The Sir William Dunn School of Pathology
South Parks Road,
Oxford OX1 3RE, UK
Tel/Fax: +44 1865 275511
quentin.sattentau@path.ox.ac.uk
The movie above represents a 3D model
of HIV-1 particles (red spheres) moving between an infected T cell (plasma
membrane in red) and an uninfected, receptor-expressing T lymphocyte (plasma
membrane in green) across a virological synapse. Tomography and modelling by Dr Sonja Welsch.
HIV-1 interactions with T cells and macrophages
Rebecca Russell, Amy Baxter and Chris Duncan
Viruses use two mechanisms to spread
within a host: release of cell-free particles and direct, cell-to-cell spread.
Cell-cell spread of HIV may confer advantages over cell-free spread, such as
more rapid viral dissemination, and may modulate certain intracellular pathways.
Using systems based on live and fixed cell imaging techniques, PCR for viral
RNA and DNA and analysis of cell activation pathways we are investigating the
molecular cell biology of HIV-1 interaction with its target cells. We coined
the term Òvirological synapseÓ in 2002, and were the
first group to demonstrate such an HIV-1-induced structure in T cells. We are
currently investigating the interactions between HIV-1-infected T cells and
macrophages. Macrophages efficiently take up HIV-1 infected T cells and degrade
them, potentially representing an innate mechanism of HIV-1 resistance.
However, during this process macrophages may become infected, thereby
generating a long-lived reservoir of HIV-1 infection. Understanding these
virus-cell interactions will yield insight into the pathogenesis of HIV-1
infection.
Funding: The MRC UK, The Wellcome Trust,
UK
Collaborations: Sonja Welsch, Heidelberg,
Germany; Kay Grunewald, STRUBI, Oxford
HIV neutralising antibody vaccine studies
Frank Wegmann, Rebecca Russell, Gemma Needham, Torben Schiffner, Leo Kong
Despite intense activity by the
international research community, we still do not have an effective vaccine
against HIV-1 infection. Neutralising antibodies are an important component of
the anti-HIV immune response and an effective vaccine will need to elicit such
antibodies. We have a growing understanding of the antigenicity
of the target of HIV-specific neutralising antibodies, the viral glycoproteins, Env. Nevertheless,
no laboratory has yet induced the requisite high levels of neutralising
antibodies following immunisation and the problem appears to be due to an
unfocussed antibody response and the innately poor immunogenicity of HIV Env. We are attempting to focus the antibody response to
the relevant neutralization epitopes on Env by a strategy of Ôimmune silencingÕ of irrelevant epitopes that has been developed by our group in
collaboration with B. Davis in the Department of Chemistry. This approach uses glycan conjugation to the antigen followed by enzymatic glycan polymerization. In a parallel study we are working
to overcome the conformational flexibility intrinsic to HIV-1 Env by using chemical cross-linking. It is hoped that this
will provide a more rigid framework for B cell recognition and help define the
parameters of B cell immunodominance related to epitope flexibility and topology. Finally we areattempting to understand how endogenous and extrinsic glycans affect the antigenicity and
immunogenicity of HIV-1 Env by structural and
molecular modelling analyses coupled with functional in vitro and in vivo
studies.
Funding: The International AIDS Vaccine Initiative, The Bill and
Melinda Gates Foundation, The James Martin 21st Century School
Vaccine Institute
Collaborations: Ben Davis and Heiko Schuster,
Department of Chemistry, University of Oxford, Chris Scanlan
and Camille Bonomelli, Department of Biochemistry, The
University of Oxford.
Novel adjuvants for systemic and
mucosal immunity
Frank Wegmann, Sarah Brinckmann, Leo Kong, Kate Gartlan
Vaccines that rely upon subunit
antigens or killed microorganisms may be lacking in immunogenicity. This is
true of HIV-1 Env, which does not carry any intrinsic
activators of innate immunity. Since HIV-1 vaccination may require immune
responses at the mucosal surfaces, we are interested in adjuvants
that may elicit immune responses both systemically and mucosally.
In this context we have discovered two adjuvants that
may have potential use in man with respect to vaccination against HIV-1 and
other pathogens. Carbopol is a polyanionic
gel that triggers a robust Th1-type adaptive immune response that is protective
when formulated with influenza HA against lethal influenza challenge in vivo,
and elicits strong antibody responses against HIV-1 Env
in various in vivo models. Polyethyleneimine (PEI) is
a cationic polymer that has strong adjuvant activity both systemically and mucosally, and drives a balanced Th1/Th2-type response in
vivo. When formulated with influenza HA or HSV gD, PEI protects against lethal challenge with
influenza and HSV respectively after in vivo immunization. Neither
adjuvant is dependent upon TLR activation for its activity, and we are
currently investigating their mechanism of action.
Funding: The Bill and Melinda Gates Foundation, EUROPRISE EU
Network of Excellence, The International AIDS Vaccine Initiative, The James Martin 21st Century School Vaccine
Institute
Collaborations: Jon Heeney, Department of
Veterinary Science, University of Cambridge
Patent filed:
PCT/GB2007/000979 - Novel adjuvant
Patent filed: GB0805356.3 – vaccine adjuvant composition
Reactive carbonyls in allergy and autoimmunity
Amin Moghaddam, Kate Gartlan, William Hillson
Reactive carbonyls
are adducted to a variety of antigens by various pathways including treatment
with aldehydes, oxidation, and by enzymatic activity.
The presence of reactive carbonyls can increase the immunogenicity of antigens,
and we have recently shown that their presence drives a Th2 bias. These
properties of carbonyls have implications for allergic and autoimmune
phenomena. We have established in vivo models based on hypersensitivity to
vaccination with Respiratory Syncytial Virus (RSV),
peanut allergy and asthma. We are also investigating the molecular mechanisms
underlying the immune modulating activity of reactive carbonyls, and are
investigating scavenger receptors and other innate immune pathways.
Collaborations: Clare Lloyd, Imperial College London
Patent:
filed ICOY/P28613GB - Reactive carbonyls as immune modulators
Additional laboratory support
á
The
laboratory is supported by The James Martin 21st Century School
Vaccine Institute
á
Q.
Sattentau is a Jenner Vaccine Institute Investigator
á
The
laboratory acknowledges support from Dormeur Investment
Service Ltd.
Ms Amy Baxter, Wellcome Trust 4 year PhD Programme in Infection and
Immunity
Ms. Sarah Brinckmann
BSc: Europrise (EU)-Funded DPhil Student
Dr. Chris Duncan MD: Wellcome Trust-funded PhD Programme for Clinicans
Dr. Kate Gartlan
BSc, PhD: EPA-funded Postdoctoral Research Scientist
Mr. William Hillson, BSc, EPA-funded DPhil probationer
Mr Leo Kong BSc: NIH-Oxford Programme DPhil Student
Dr. Amin
Moghaddam MD, MSc: Senior Postdoctoral Research Scientist
Ms Gamma Needham BSc: Research
Assistant and Assistant Laboratory Manager
Dr. Rebecca Russell BSc, PhD:
IAVI-funded Postdoctoral Research Scientist and Laboratory Manager
Dr. Quentin Sattentau, BSc, PhD: Laboratory PI
Mr. Torben
Schiffner, BSc: Departmental DPhil student
Dr. Frank Wegmann,
BSc, PhD: Gates Foundation Grand Challenge Postdoctoral Scientist
Co-supervised staff and students
Dr. Catherine Sargent MD: MRC-funded DPhil Student with Prof. Sir Andrew
McMichael, WIMM, University of Oxford
Dr. Chris Duncan MD: Wellcome Trust-funded PhD Programme for Clinicans
with Prof. Sir Andrew McMichael, WIMM, University of
Oxford
Previous members of the Oxford laboratory
Dr. Katherine Gantlett (nŽe
Fowler), Wellcome Trust Infection and Immunity Dphil (2002-2005): now
Vaccine Clinical trials manager, The Jenner Vaccine Institute, University of
Oxford
Dr. Emma Jones, Dphil, EU-funded
Postdoctoral Research Scientist (2005-2006):
now Postdoctoral Research Staff, Cardiff University, Wales
Dr. Neil Sheppard, BSc, MRC-funded DPhil student
(2003-2006), Gates Foundation-funded Postdoctoral Scientist (2006-2007): now Senior Scientist, Vaccine Discovery Group,
Pfizer La Jolla, CA USA
Dr. Samer Sourial,
BSc, PhD, Postdoctoral Research Scientist (2007-2008): now Graduate-entry Medical Student,
Liverpool UK
Dr. Ivonne Mitar,
BSc, DPhil, MRC-funded Research Assistant and Dphil
student (2002-2007): now Staff member,
Health Interactions, Scientific Consultancy, UK
Dr. Daniela Romer, BSc, MSc, DPhil, EU-funded DPhil student (2003-2007): now Medical Department of Sanofi-Aventis Germany
Ms. Cynthia Robson, BSc, Laboratory Manager and Graduate
Research Assistant (2005-2007): now Funding
Administrator, Auckland University
Dr. Clare Jolly, PhD, Wellcome
Trust then MRC-funded Postdoctoral Research Scientist (2000-2008): now MRC Career Development Award Fellow,
UCL London UK
Dr. Sinead Brady, PhD, IAVI-funded Postdoctoral Research
Scientist (2007-2009): now
Dr. Giulia Zanetti, Wellcome Trust funded DPhil (2006-2009): now Postdoctoral Scientist, University of Berkley, USA
Dr. Stefanie Michor, Bill and
Melinda Gates Foundation funded DPhil (2005-2009): now
Dr. Nicola Martin BM, DPhil, MRC-funded Dphil
student (2006-2009): now clinical school
medicine, University of Oxford
Dr. Fedde Groot BSc, PhD, amFAR Postdoctoral Research Fellow (2007-2009): now Graduate-entry medicine, University of
Oxford
Ms. Karoliina Laamanen BSc, Bill and Melinda
Gates Foundation-funded Graduate Research Assistant (2006-2010): now Senior Technical Assistant at the
Helsinki Blood Sevice, Finland
Dr. George Krashias BSc, MSc, Bodossaki Foundation DPhil Student
(2006-2009): now Lecturer, University of
Nicosia, Cyprus
Selected recent publications (from a total of 140)
11. Montefiori D, Sattentau QJ, Flores J, Esparza J, Mascola J, Working group convened by the Global HIV vaccine Enterprise (2008). Antibody-based HIV-1 vaccines: recent developments and future directions. Plos Medicine 4: e348.
23. Mitchison NA, Sattentau
QJ. (2005). Fundamental Immunology and What it Can
Teach us About HIV Vaccine Development. Curr Drug
Targets Infect Disord. 5:87-93.
24. Sheppard
N, Sattentau QJ. (2005). The prospects for vaccines against HIV-1: more than a
field of long-term non-progression? Expert Rev Mol Med. 7:1-21.
25. Jolly, C and
Sattentau, QJ. (2004). Retroviral Spread by Induction of Virological
Synapses. Traffic 5:
643—650.
26. Piguet, V and
Sattentau, QJ. (2004). Dangerous Liasons at the Virological Synapse. J. Clin. Invest.
114: 2-8.